articles | articles accepted | book chapters


2011 Souza E D S, Willemart R H. Harvest-ironman: heavy armature, and not its defensive secretions, protects a harvestman against a spider. Animal Behaviour, 81: 127-133. doi:10.1016/j.anbehav.2010.09.023 PDF

2011 Willemart R H, Santer R D, Spence A J, Hebets E A. A sticky situation: solifugids (Arachnida, Solifugae) use adhesive organs on their pedipalps for prey capture. Journal of Ethology, 29: 177-180. doi:10.1007/s10164-010-0222-4 PDF

2010 Willemart R H, Giribet G. A scanning electron microscopic survey of the cuticle in Cyphophthalmi (Arachnida, Opiliones) with the description of novel sensory and glandular structures. Zoomorphology, 129: 175-183. doi:10.1007/s00435-010-0110-z PDF

2010 Willemart R H, Pérez-González A, Farine J, Gnaspini P. Sexually dimorphic tegumental gland openings in Laniatores (Arachnida, Opiliones), with new data on 23 species. Journal of morphology, 271: 641-653. doi:10.1002/jmor.10822 pmid:20027634 PDF

2009 Chelini M, Willemart R H, Hebets E A. Costs and benefits of freezing behaviour in the harvestman Eumesosoma roeweri (Arachnida, Opiliones). Behavioural processes, 82: 153-159. doi:10.1016/j.beproc.2009.06.001 pmid:19539731 PDF

2009 Willemart R H, Farine J, Gnaspini P. Sensory biology of Phalangida harvestmen (Arachnida, Opiliones): a review, with new morphological data on 18 species. Acta Zoologica, 90: 209-227. doi:10.1111/j.1463-6395.2008.00341.x PDF

2009 Willemart R H, Osses F, Chelini M, Macías-Ordóñez R, Machado G. Sexually dimorphic legs in a neotropical harvestman (Arachnida, Opiliones): ornament or weapon?. Behavioural processes, 80: 51-59. doi:10.1016/j.beproc.2008.09.006 pmid:18929628 PDF

2007 Peretti A V, Willemart R H. Sexual coercion does not exclude luring behavior in the climbing camel-spider Oltacola chacoensis (Arachnida, Solifugae, Ammotrechidae). Journal of Ethology, 25: 29-39. doi:10.1007/s10164-006-0201-y PDF

2007 Willemart R H, Chelini M. Experimental demonstration of close-range olfaction and contact chemoreception in the Brazilian harvestman, Iporangaia pustulosa. Entomologia Experimentalis et Applicata, 123: 73-79. doi:10.1111/j.1570-7458.2007.0527.x PDF

2007 Willemart R H, Chelini M, de Andrade R, Gnaspini P. An ethological approach to a SEM survey on sensory structures and tegumental gland openings of two neotropical harvestmen (Arachnida, Opiliones, Gonyleptidae). Italian Journal of Zoology, 74(1): 39-54. doi:10.1080/11250000601090529 PDF

2006 Willemart R H, Pellegatti-Franco F. The Spider Enoploctenus Cyclothorax (Araneae, Ctenidae) Avoids Preying on the Harvestman Mischonyx Cuspidatus (Opiliones, Gonyleptidae). Journal of Arachnology, 34: 649-652. doi:10.1636/S05-70.1 PDF

2006 Willemart R H, Farine J, Peretti A V, Gnaspini P. Behavioral roles of the sexually dimorphic structures in the male harvestman, Phalangium opilio (Opiliones, Phalangiidae). Canadian Journal of Zoology, 84: 1763-1774. doi:10.1139/Z06-173 PDF

2004 Willemart R H, Gnaspini P. Spatial distribution, mobility, gregariousness, and defensive behaviour in a Brazilian cave harvestman Goniosoma albiscriptum (Arachnida, Opiliones, Gonyleptidae). Animal Biology, 54(3): 221-235. doi:10.1163/1570756042484674 PDF

2004 Willemart R H, Gnaspini P. Breeding biology of the cavernicolous harvestman Goniosoma albiscriptum (Arachnida, Opiliones, Laniatores): sites of oviposition, egg batches characteristics and subsocial behaviour. Invertebrate Reproduction and Development, 45(1): 15-28. PDF

2004 Willemart R H, Kaneto G E. On the natural history of the Neotropical spider Enoploctenus cyclothorax (Araneae, Ctenidae). Bulletin of the British Arachnological Society, 13(2): 53-59. PDF

2003 Willemart R H, Gnaspini P. Comparative Density of Hair Sensilla on the Legs of Cavernicolous and Epigean Harvestmen (Arachnida: Opiliones). Zoologischer Anzeiger – A Journal of Comparative Zoology, 242: 353-365. doi:10.1078/0044-5231-00109 PDF

2002 Willemart R H. Cases of intra- and inter-specific food competition among Brazilian harvestmen, in captivity (Opiliones, Laniatores, Gonyleptidae). Revue Arachnologique 14: 49-58.

2001 Willemart R H. Egg Covering Behavior of the Neotropical Harvestman Promitobates Ornatus (Opiliones, Gonyleptidae). Journal of Arachnology, 29: 249-252. doi:10.1636/0161-8202(2001)029[0249:ECBOTN]2.0.CO;2 PDF

Articles accepted

Chelini M C, Willemart R H, Gnaspini P. Caves as a Winter Refuge by a Neotropical Harvestman (Arachnida, Opiliones). Journal of Insect Behavior

Willemart R H, Hebets E A. Sexual differences in the behavior of the harvestman Leiobunum vittatum (Opiliones, Sclerosomatidae) towards conspecific cues. Journal of Insect Behavior

Book chapters

2007 Willemart R H. Rearing and Maintenance of Animals in Captivity. Em: Harvestmen: the biology of Opiliones (Ed. Pinto-da-Rocha R, Machado G, Giribet G), Cambridge, Harvard University Press, pp: 520-524.

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